The biggest protests by Tibetan monks in nearly two decades have rippled into Chinese provinces populated by Tibetans, as the government's tough response draws condemnation from international groups.
The demonstrations over past days have followed marches around the world to mark the 49th anniversary of an uprising against Communist rule in the remote, mountainous region that has become a focus for protest ahead of this year's Beijing Olympics.
While China has focused on condemning foreign-based critics of its presence in Tibet, the shows of bold defiance within its borders are likely to make security preparations for the Olympic Games an even bigger worry for officials.
"The reports of protests outside Lhasa show that Tibetans know the eyes of the world are upon them and are determined not to let the momentum drop," Matt Whitticase of the London-based Free Tibet Campaign said.
On Monday, 500 monks from Drepung monastery defied authorities by staging a rare march in the Tibetan capital of Lhasa, an act that the Chinese government called "an illegal activity that threatened social stability".
About 2000 Chinese security personnel fired tear gas to try to disperse 600 monks from Sera monastery taking part in a second day of street protests in Lhasa, a source said.
They demanded the release of about a dozen fellow monks from Sera detained this month for waving a Tibetan flag and shouting pro-independence slogans, the source said.
"The demonstrations are the largest by monks since the 1989 protests that led to the imposition of martial law in Tibet's capital," the International Campaign for Tibet said in a statement.
Another rights group said about 400 monks from Lutsang monastery in the northwestern province of Qinghai, known in Tibetan as Amdo, protested on Monday and shouted slogans for their exiled spiritual leader, the Dalai Lama, to return.
The Dalai Lama fled to India after the failed uprising in 1959, nine years after People's Liberation Army troops marched into the predominantly Buddhist Himalayan region. The protesters shouted "Free Tibet!", the Free Tibet Campaign said on Thursday.
About 100 monks from Myera monastery in the neighbouring province of Gansu also protested on Monday, the rights group said, adding that police were investigating who was involved.
A source with knowledge of the protests quoted monks and witnesses as saying the sound of gunfire was heard outside the walls of monasteries. But no casualties have been reported.
Chinese Foreign Ministry spokesman Qin Gang said the protesters in Lhasa were "seeking to spark social turmoil".
"This was carefully planned by the Dalai clique in a bid to separate Tibet and sabotage Tibetan people's normal life of stability and harmony," he told a regular news conference.
The strife "could be a harbinger of further clashes between Tibetans and Chinese authorities in this Olympics year", said Mary Beth Markey, vice president of the International Campaign for Tibet.
The demonstrations in the heavily policed region are precisely what China's Communist leaders are keen to avoid ahead of the Olympics starting Aug 8.
On Wednesday, China closed the north face of Mount Everest to expeditions until after the Olympic torch ascends the peak in early May. Five Americans, including an ethnic Tibetan, unfurled "Free Tibet" banners on Everest – known in China by its Tibetan name, Qomalangma – last year.
China's neighbour, India, which hosts many exiled Tibetans, has been careful to distance itself from the protests.
Indian police arrested around 100 Tibetans on Thursday, dragging them into police vans, when they tried to march to the Chinese border to press claims for independence and protest the Olympics.
The marchers set off on Monday as part of the global protests, leaving from Dharamsala, home to the Dalai Lama and the refugees' "government-in-exile".
The Indian police have said they are acting on government orders to restrain the marchers, claiming they have breached an agreement not to hold "anti-Chinese activities" on Indian soil.
Asked if China was satisfied with India's handling of the protests, Qin held back from direct comment.
"We hope that the Indian side will, based on broader considerations of bilateral relations, abide by the promise it made," he said.